Six New Papers from the Forthcoming Volume 'Anatomy, Phylogeny, and Palaeobiology of Early Archosaurs and their Kin.

You can access the abstracts electronically here.

Mastrantonio, B. M., Schultz, C. L., Desojo, J. B., and J.  Bittencourt Garcia. 2013. The braincase of Prestosuchus chiniquensis (Archosauria: Suchia)From: Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds) 2013. Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin. Geological Society, London, Special Publications, 379, doi:10.1144/SP379.10

Abstract - The osteology of an almost complete braincase of the rauisuchian archosaurs Prestosuchus chiniquensis from the Middle Triassic of Brazil is described for first time, based on two specimens (UFRGS-PV-0629-T and UFRGS-PV-0156-T). A comparative description with other taxa of rauisuchians is presented that forms the basis of a phylogenetic analysis. To perform the phylogenetic analysis, we describe and discuss each character codification for a modified version of the recent matrices of Gower (2002), Gower & Nesbitt (2006) and Brusatte et al.(2010). The analysis resulted in two most parsimonious trees that differ from the topologies recovered by Gower (2002) in a few aspects within Rauisuchia, and Prestosuchus chiniquensis was unequivocally depicted as deeply nested within Pseudosuchia, as the sister taxon of Batrachotomus kuperferzellensis in both topologies, supported by a single synapomorphy: the reduced to small fissure of the post-temporal fenestra between parietal, supraoccipital and exoccipital-opisthotic.


Sues, H.-D., Desojo, J. B., and M. D. Ezcurra. 2013. Doswelliidae: a clade of unusual armoured archosauriforms from the Middle and Late Triassic. From: Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds) 2013. Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin. Geological Society, London, Special Publications, 379, first published on April 23, 2013, doi:10.1144/SP379.13

Abstract - Doswelliidae is a clade of armoured non-archosaurian archosauriform reptiles more closely related to Archosauria than are Proterosuchidae, Erythrosuchidae and possibly Euparkeria capensis. It is currently known from the late Middle Triassic (Ladinian) of Germany, the late Middle to early Late Triassic (Ladinian–Carnian) of Argentina and Brazil, and the Late Triassic (Carnian–Norian) of the USA. To date, two unambiguous synapomorphies diagnose Doswelliidae: (i) osteoderm ornamentation coarse, incised, and composed of central regular pits of subequal size and shape, and (ii) osteoderms with anterior articular lamina. Five taxa are currently recognized: Archeopelta arborensis, Doswellia kaltenbachi, Doswellia sixmilensis, Tarjadia ruthae and a new taxon from Germany. Based on skeletal features and occurrence, doswelliid archosauriforms may have had a semi-aquatic mode of life.


Langer, M. C., and J. Ferigolo. 2013. The Late Triassic dinosauromorph Sacisaurus agudoensis (Caturrita Formation; Rio Grande do Sul, Brazil): anatomy and affinities. From: Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds) 2013. Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin.  Geological Society, London, Special Publications, 379, first published on April 23, 2013, doi:10.1144/SP379.16

Abstract - Silesauridae is an exclusively Triassic group of dinosauromorphs, knowledge on the diversity of which has increased dramatically in the last few years. Silesaurid relationships are still contentious, as a result in part of different homology statements, particularly regarding the typical edentulous mandible tip of these animals. One of the most complete silesaurids yet discovered is Sacisaurus agudoensis from the Caturrita Formation (Late Triassic: Norian) of Rio Grande do Sul, Brazil, represented by numerous isolated bones recovered from a single site. The anatomy of S. agudoensis is fully described for the first time here, and comparisons are provided to other basal dinosauromorphs. S. agudoensis is a small-bodied animal (less than 1 m in length) that possesses a dentition consisting of leaf-shaped crowns with large denticles in the carinae, a plesiomorphic propubic pelvis with an almost fully closed acetabulum, elongate distal hindlimbs suggesting well-developed cursorial ability, and a laterally projected outer malleolus in the tibia. All previous numerical phylogenies supported a non-dinosaur dinosauromorph affinity for Silesauridae, but the reanalysis of one of those studies suggests that a position within Dinosauria is not unlikely, with silesaurids forming the basal branch of the ornithischian lineage.


Turner, A. H., and S. J. Nesbitt. 2013. Body size evolution during the Triassic archosauriform radiation. From: Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds) 2013. Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin.  Geological Society, London, Special Publications, 379, first published on April 23, 2013, doi:10.1144/SP379.15

Abstract - The first large (>1 m) diapsids appeared near the Permian–Triassic extinction and a subset of diapsids, the archosauriforms, expanded their body size range soon after in the Early–Middle Triassic. Here, we examine body size at key evolutionary events within Archosauriformes during the Triassic and through the end-Triassic extinction. Using femoral length as a body size proxy and a temporally calibrated phylogeny of Archosauriformes, we estimate ancestral body sizes using a maximum likelihood approach and test for the presence of an adapative radiation by comparing the fit of competing evolutionary models. Archosauriform body size is characterized by punctuated change with more change occurring early in the Triassic. Archosaurs crossing the Triassic–Jurassic boundary show a wide range in ancestral size, and dinosaurs (sauropodomorphs and theropods) are considerably larger in the Jurassic. Crocodylomorph origins are characterized by a drop in body size; however, both the relative amount of change and the rate of change are matched among other archosaur clades. Archosauriforms increase in absolute body size through the Triassic and evidence suggests that a directional trend in size increase occurred in the early Mesozoic. The morphological signature of adaptive radiation is rare in comparative data from extant animals but is present at the origination of Archosauriformes.


Niedźwiedzki, G., Brusatte, S. L., and R. J. Butler. 2013. Prorotodactylus and Rotodactylus tracks: an ichnological record of dinosauromorphs from the Early–Middle Triassic of Poland. From: Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds) 2013. Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin. Geological Society, London, Special Publications, 379, first published on April 23, 2013, doi:10.1144/SP379.12

Abstract - We present the first comprehensive description of Prorotodactylus and Rotodactylus  dinosauromorph tracks from the Early and Middle Triassic of the Holy Cross Mountains, Poland. We describe and comprehensively figure tracks that have been mentioned briefly in previous accounts as well as new, recently discovered material, and analyse the variation and stratigraphic distribution of these specimens. Tracks have been recorded from four sites – Koszary, Stryczowice, Wióry and Baranów – which span the early Olenekian – early Anisian (c.250–246 Ma). These tracks therefore represent an ichnological record of the evolutionary succession of early dinosauromorphs during the earliest part of their evolutionary history. Recognized track types include cf. Prorotodactylus isp., Prorotodactylus isp., Prorotodactylus mirus, Rotodactylus cursorius, Rotodactylus isp. and cf. Rotodactylus isp. At least three distinct Early and early Middle Triassic early dinosauromorph ichnofaunas can be recognized. The oldest, which is early Olenekian in age, is characterized by the presence of Prorotodactylus isp., cf.Prorotodactylus isp. and non-archosaurian archosauromorph or archosaur tracks (e.g. Synaptichnium isp., Protochirotherium isp.), recorded at the Stryczowice and Koszary sites. The following assemblage, recorded at the late Olenekian Wióry site, displays the highest ichnodiversity of dinosauromorphs, with four track types present (Prorotodactylus isp., Prorotodactylus mirus, Rotodactylus cursorius and cf. Rotodactylus isp.). The youngest site, Baranów, includes Rotodactylus isp., as well as other larger dinosauromorph tracks. The first body fossil evidence of dinosauromorphs is a few million years younger than the youngest Polish tracks, so Prorotodactylus and Rotodactylus tracks currently provide the oldest record of dinosauromorph morphology, biology and evolution.


Ezcurra, M. D., Butler, R. J., and D. J. Gower. 2013. ‘Proterosuchia’: the origin and early history of Archosauriformes.. From: Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds) 2013. Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin. Geological Society, London, Special Publications, 379, first published on April 23, 2013, doi:10.1144/SP379.11

Abstract - The earliest history of Archosauriformes is mainly represented by members of Proterosuchidae and Erythrosuchidae, which are known worldwide from latest Permian to Middle Triassic beds. These two groups were historically combined within ‘Proterosuchia’, with approximately 30 nominal species. Two morphotypes have been recognized among proterosuchians: proterosuchids with a generally more sprawling gait and elongated and low skulls with an overhanging premaxilla, and the more heavily built erythrosuchids, with a probably less sprawling gait and large, presumably hypercarnivorous, skulls. The systematics of ‘Proterosuchia’ was relatively chaotic throughout most of the twentieth century, but currently there exists consensus regarding the non-monophyly of proterosuchians and their phylogenetic position outside all other archosauriforms. In contrast, the delimitation and taxonomic content of Proterosuchidae and Erythrosuchidae remain unstable. Few studies of proterosuchian palaeobiology have been carried out. Current lines of evidence favour a predominantly terrestrial lifestyle for proterosuchians. Limb bone histology indicates rapid continuous growth rates in Proterosuchus and Erythrosuchus before reaching sexual maturity. A better knowledge of proterosuchian anatomy, systematics, evolution and ecology is important for advancing understanding of the origin and early radiation of Archosauriformes and the patterns of biotic recovery following the Permo-Triassic mass extinction event. There remains much research to be carried out in proterosuchian palaeobiology.

Two New Triassic Archosaurian Taxa in the New Issue of Acta Palaeontologica Polonica

These two papers have been available for awhile as accepted manuscripts, but now they are officially out and freely downloadable.

Niedźwiedzki, G., Sulej, T., and J. Dzik. 2012. A large predatory archosaur from the Late Triassic of Poland. Acta Palaeontologica Polonica 57 (2): 267-276. doi: http://dx.doi.org/10.4202/app.2010.0045

Abstract - We describe a new large predatory archosaur, Smok wawelski gen. et sp. nov., from the latest Triassic (latest Norian–early Rhaetian; approximately 205–200 Ma) of Lisowice (Lipie Śląskie clay−pit) in southern Poland. The length of the reconstructed skeleton is 5–6 m and that of the skull 50–60 cm, making S. wawelski larger than any other known predatory archosaur from the Late Triassic and Early Jurassic of central Europe (including theropod dinosaurs and “rauisuchian” crurotarsans). The holotype braincase is associated with skull, pelvic and isolated limb−bones found in close proximity (within 30 m), and we regard them as belonging to the same individual. Large, apparently tridactyl tracks that occur in the same rock unit may have been left by animals of the same species. The highly autapomorphic braincase shows large attachment areas for hypertrophied protractor pterygoideus muscles on the lateral surface and a wide, funnel−like region between the basal tubera and basipterygoid processes on the ventral surface. The skeleton (cranial and postcranial) possesses some features similar to those in theropod dinosaurs and others to those in large crocodile−line archosaurs (“rauisuchians”), rendering phylogenetic placement of S. wawelski difficult at this time.

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Kammerer, C. F., Nesbitt, S. J., and N. H. Shubin. 2012. The first silesaurid dinosauriform from the Late Triassic of Morocco. Acta Palaeontologica Polonica 57 (2): 277-284 doi: http://dx.doi.org/10.4202/app.2011.0015

Abstract - Disarticulated material from the Late Triassic Timezgadiouine Formation in the Argana Basin of Morocco represents a new taxon of silesaurid dinosauromorph, Diodorus scytobrachion gen. et sp. nov. D. scytobrachion can be distinguished from other silesaurids by the presence of anteriorly−canted teeth that decrease in size towards the anterior end of the dentary and a distinct lateral ridge running parallel to the dentary alveolar margin. In a phylogenetic analysis, D. scytobrachion is recovered as the sister−taxon to the Brazilian Sacisaurus agudoensis, nested deep within Silesauridae. This new taxon provides further evidence of a near−cosmopolitan range for basal dinosauriforms in the Late Triassic and further demonstrates the disparity of dental morphologies within Silesauridae.

Evolution of Bipedality and Herbivory Among Triassic Dinosauromorphs

Note that this paper is in Japanese with an English version of the abstract.

Kubo, T. 2011. Evolution of bipedality and herbivory among Triassic dinosauromorphs. Memoir of the Fukui Prefectural Dinosaur Museum 10:55－62．

Abstract - Discoveries of Triassic non-dinosaur dinosauromorphs since 2000 revealed that they were more widely spread chronologically and geographically than previously thought. A member of silesaurids, the sister clade of dinosaurs, Silesaurus was a quadrupedal and herbivorous animal that differs considerably from the condition previously assumed for the ancestor of dinosaurs that are bipedal and carnivorous. Currently, stance and diet of the common ancestor of dinosaurs are not clear. To redeem this situation, Ancestral State Reconstruction methods were conducted to infer how quadrupedality and herbivory were evolved among dinosauromorphs. The results of analyses indicate that quadrupedal stance evolved only among silesaurids. Herbivorous diet was readily evolved from carnivorous diet among Dinosauromorpha and the ancestral state reconstruction using likelihood methods indicated that the possibility of the common ancestor of dinosaurs being herbivore is more than 60%.

Diodorus scytobrachion, A New Silesaurid from the Late Triassic of Morocco

I've had this paper in the queue for a bit as I still don't like the idea of online publishing of new taxon names at the "accepted manuscript" stage, but the knowledge of this proposed new taxon is increasingly widespead so I'll post the abstract and link here now. Silesaurids are amazingly cosmopolitan in their distribution and must have been significant members of various Triassic ecosystems, most likely filling the role later occupied by ornithischians.

I'm a big fan of the creative taxonomic name as well.

Kammerer, C. F., Nesbitt, S. J., and N. H. Shubin. In press. The first basal dinosauriform (Silesauridae) from the Late Triassic of Morocco. Acta Palaeontologica Polonica. doi: 10.4202/app.2011.0015

Abstract - Disarticulated material from the Late Triassic Timezgadiouine Formation in the Argana Basin of Morocco represents a new taxon of silesaurid dinosauromorph,

Diodorus scytobrachion gen. et sp. nov. D. scytobrachion can be distinguished from other silesaurids by the presence of anteriorly-canted teeth that decrease in size towards the anterior end of the dentary and a distinct lateral ridge running parallel to the dentary alveolar margin. In a phylogenetic analysis, D. scytobrachion is recovered as the sister taxon to the Brazilian Sacisaurus agudoensis, nested deep within Silesauridae. This new taxon provides further evidence of a near-cosmopolitan range for basal dinosauriforms in the Late Triassic and further demonstrates the disparity of dental morphologies within Silesauridae.

New Information on the Triassic Dinosauriform Silesaurus opolensis

Piechowski, R., and J. Dzik. 2010. The axial skeleton of Silesaurus opolensis. Journal of Vertebrate Paleontology 30:1127 – 1141. DOI: 10.1080/02724634.2010.483547



Abstract - A recent find of an articulated skeleton of Silesaurus opolensis at its early Late Triassic type locality Krasiejów (Poland), with skull, neck, pectoral girdle, and thorax, supplemented by additional preparation of previously collected articulated specimens, enables complete restoration of the vertebral column and associated skeletal parts. Cervical ribs of Silesaurus, well preserved in their original disposition, are parallel to the neck and extend backward for a few vertebral lengths. There is a sudden change in their morphology behind the seventh vertebra, although otherwise the transition from the cervical to the dorsal vertebrae is very gradual. Parapophyses slowly migrate upward along the anterior margin of the centrum and leave the centrum at the sixth or seventh dorsal vertebra. Narrowing of the dorsal extremities of the neural spines of the fourth and neighboring vertebrae suggests the ability of this region of the vertebral column to bent upward. There is thus a disparity between the structural and functional neck-thorax transition. The presence of three sacrals firmly connected by their ribs with the ilia and the long tail of Silesaurus, providing a counterbalance to the weight of the body in front of the pelvis, suggests the ability for fast bipedal running. However, unusually long but gracile forelimbs of Silesaurus suggest that it represents a transition towards secondarily quadrupedal locomotion, characterizing most of the later herbivorous dinosaurs.

Asilisaurus kongwe, the Oldest Avian-line Archosaur and the Early Diversification of Ornithodira

Nesbitt, S.J., Sidor, C.A., Irmis, R.B., Angielczyk, K.D., Smith, R.M.H., and L. A. Tsuji. 2010. Ecologically distinct dinosaurian sister group shows early diversification of Ornithodira. Nature. doi:10.1038/nature08718

Abstract - The early evolutionary history of Ornithodira (avian-line archosaurs) has hitherto been documented by incomplete (Lagerpeton) or unusually specialized forms (pterosaurs and Silesaurus). Recently, a variety of Silesaurus-like taxa have been reported from the Triassic Period of both Gondwana and Laurasia, but their relationships to each other and to dinosaurs remain a subject of debate. Here we report on a new avian-line archosaur from the early Middle Triassic (Anisian) of Tanzania. Phylogenetic analysis places Asilisaurus kongwe gen. et sp. nov. as an avian-line archosaur and a member of the Silesauridae, which is here considered the sister taxon to Dinosauria. Silesaurids were diverse and had a wide distribution by the Late Triassic, with a novel ornithodiran bauplan including leaf-shaped teeth, a beak-like lower jaw, long, gracile limbs, and a quadrupedal stance. Our analysis suggests that the dentition and diet of silesaurids, ornithischians and sauropodomorphs evolved independently from a plesiomorphic carnivorous form. As the oldest avian-line archosaur, Asilisaurus demonstrates the antiquity of both Ornithodira and the dinosaurian lineage. The initial diversification of Archosauria, previously documented by crocodilian-line archosaurs in the Anisian, can now be shown to include a contemporaneous avian-line radiation. The unparalleled taxonomic diversity of the Manda archosaur assemblage indicates that archosaur diversification was well underway by the Middle Triassic or earlier.

Life restoration of Asilisaurus with sail-backed poposauroid in the background. Image by M.H. Donnelly (Field Museum).

Prior to 2003 the non-dinosaurian dinosauriforms known as silesaurids were unrecognized in the fossil record. Specimens existed in collections, collected as early as the 1930s, while others were given tentative identifications (e.g., the ornithosuchid of Long and Murry, 1995 and the ornithischian Technosaurus). Dzik (2003) described the first, Silesaurus opolensis from the Carnian of Poland, with its very distinctive femoral and morphologies. Suddenly similar forms were recognized from all over the globe (e.g., Eucoelophysis, Sacisaurus; Ezcurra 2006; Nesbitt et al. 2007; Irmis et al. 2007a), whereas new specimens were being discovered from the Chinle Formation of New Mexico and Arizona (Parker et al. 2006; Irmis et al. 2007b).

Still, because the earliest pseudosuchian archosaurs were known from the Anisian (e.g., the Moenkopi Formation of Arizona), whereas the earliest ornithodirans were from the Ladinian of Argentina, there was a proposed ghost lineage for Ornithodira existing back into the Anisian.

Sterling Nesbitt looks over the Ruhuhu Valley in 2007. Photo by L. Tsuji.

One of the strengths of phylogentic analysis is the ability to make predictions as to where in time and space certain groups should and could be found. Sterling Nesbitt clearly recognized the strong possibility that the earliest representatives of ornithodira could be found in the Manda beds of Tanzania as this fauna was known for its pseudosuchian constituents. A few years back this prediction paid off, as a team lead by Christian Sidor in 2007 to explore the Permian and Triassic rocks of Tanzanian, uncovered an amazing deposit of early archosaurs including numerous specimens of a silesaurid. Named Asilisaurus kongwe (Ancient ancestor lizard) these specimens represent the earliest known member of the lineage leading to dinosaurs and strongly supports the diversification of the Archosauria by the early Middle Triassic (~243 million years ago).

The tibia of Asilisaurus, following excavation in 2007. Photo by R. Smith.

This find also strongly suggests that adaptations for an omnivorous or herbivorous diet evolved independently in silesaurids, ornithischians, and sauropodomorphs, from carnivorous ancestors. The paper also phylogenetically defines the Silesauridae and proposes that the South American archosaurs Lewisuchus and Pseudlagosuchus are members of this clade and probably also synonymous. Another early appearance of a non-dinosaurian dinosauriform in
Gondwana also provides further support for a southern origin for this group.

Skeletal reconstruction of Asilisaurus, with missing bones in gray. Image by S. Nesbitt.

One final point, silesaurs have been proposed to be basal ornithischians (e.g., Ferigolo & Langer, 2007) and represent the Late Triassic radiation of that clade. However, if this is the case, despite the lack of support in phylogentic analyses (e.g., Irmis et al., 2007b), the discovery of early Middle Triassic ornithichians would pull the split with saurischians back as early as the Early Triassic and create a sizeable ghost lineage for the base of Saurischia.

More photos and information on Ruhuhu Basin Research can be found here:
http://protist.biology.washington.edu/sidor/Ruhuhu/asilisaurus.html

REFERENCES

Dzik, J. 2003. A beaked herbivorous archosaur with dinosaur affinities from the early Late Triassic of Poland. Journal of Vertebrate Paleontology 23:556-574.

Ezcurra, M.D. 2006. A review of the systematic position of the dinosauriform archosaur Eucoelophysis baldwini from the Upper Triassic of New Mexico, U.S.A. Geodiversitas 28:649-684.

Ferigolo, J. and M. Langer. 2007. A late Triassic dinosauriform from south Brazil and the origin of the ornithischian predentary bone. Historical Biology 19:23-33.

Irmis, R.B., Parker, W.G., Nesbitt, S.J., and J. Liu. 2007a. Early ornithischian dinosaurs: the Triassic record. Historical Biology 19:3-22.

Irmis, R. B., Nesbitt, S. J., Padian, K., Smith, N. D., Turner, A. H., Woody, D., and A. Downs. 2007a. A Late Triassic dinosauromorph assemblage from New Mexico and the rise of dinosaurs. Science 317:358-361.

Nesbitt, S.J., Irmis, R.B., and W.G. Parker. 2007. A critical re-evaluation of the Late Triassic dinosaur taxa of North America. Journal of Systematic Palaeontology 5:209-243.

Parker, W.G., Irmis, R.B., and S.J. Nesbitt. 2006. Review of the Late Triassic dinosaur record from Petrified Forest National Park, Arizona. Museum of Northern Arizona Bulletin 62:160-161.

Defining Silesauridae

The new paper by Max Langer and collaborators (Langer et al., 2009) provides a phylogenetic definition for the clade Silesauridae. Silesaurids are gracile, quadrupedal, hebivorous/omnivorous dinosauriforms whose first announcement in 2003 (Dzik, 2003) generated a large amount of interest in early dinosaur phylogeny. In fact there were at least five presentations at this years SVP meeting alone that dealt with silesaurs.

Langer et al. (2009) define Silesauridae as "all archosaurs closer to Silesaurus opolensis, than to Heterodontosaurus tucki and Marasuchus lilloensis". Currently it is pretty much agreed that Silesauridae contains Silesaurus opolensis, Sacisaurus agudoensis, Eucoelophysis baldwini, and Pseudolagosuchus major (Nesbitt et al., 2005; Ezcurra, 2006; Irmis et al., 2007b; Langer et al., 2009). Silesaurus also possibly contains Lewisuchus admixtus, Technosaurus smalli, and an isolated specimen from Petrified Forest National Park (Irmis et al., 2005; Parker et al., 2006; Nesbitt et al., 2007; Langer et al., 2009).

Since their first publication it has been debated whether this clade represent dinosauriforms (e.g., Parker et al., 2006; Irmis et al., 2007a; Nesbitt et al., 2007) or possibly basal ornithischians (e.g., Ferigolo and Langer, 2007; Dzik and Sulej, 2007). Currently there is much more support for placement as the sister taxon of Dinosauria (Ezcurra, 2006; Irmis et al. 2007b, Langer et al., 2009). Nonetheless, silesaurids most likely filled an ecological niche later filled by ornithischian and thus possibly supressed early ornithischian diversity through the Late Triassic.

REFERENCES

Dzik, J. 2003. A beaked herbivorous archosaur with dinosaur affinities from the early Late Triassic of Poland. Journal of Vertebrate Paleontology 23:556-574.

Dzik, J., and T. Sulej. 2007. A review of the Early Triassic Krasiejow biota from Silesia, Poland. Paleontologia Polonica 64:3-27.

Ezcurra, M. D. 2006. A review of the systematic position of the dinosauriform archosaur Eucoelophysis baldwini from the Upper Triassic of New Mexico, U.S.A. Geodiversitas 28:649-684.

Ferigolo, J. and M. Langer. 2007. A late Triassic dinosauriform from south Brazil and the origin of the ornithischian predentary bone. Historical Biology 19:23-33.

Irmis, R. B., Parker, W. G., and S. N. Nesbitt, 2005. Critical review of the Late Triassic dinosaur record, part 2: Ornithischia. Society of Vertebrate Paleontology 25(3):73A.

Irmis, R. B., Parker, W. G., Nesbitt, S. J., and J. Liu, 2007. Early ornithischian dinosaurs: the Triassic Record. Historical Biology 19:3-22.

Irmis, R. B., Nesbitt, S. J., Padian, K., Smith, N. D., Turner, A. H., Woody, D., and A. Downs. 2007. A Late Triassic dinosauromorph assemblage from New Mexico and the rise of dinosaurs. Science 317:358-361.

Nesbitt, S. N., Irmis, R. B., and W. G. Parker, 2005. Critical review of the Late Triassic dinosaur record, part 3: Saurischians of North America. Society of Vertebrate Paleontology 25(3):96A.

Nesbitt, S. J., Irmis, R. B., and W. G. Parker. 2007. A critical reevaluation of the Late Triassic dinosaur taxa of North America. Journal of Systematic Palaeontology 5:209-243.

Parker, W. G., Irmis, R. B., and S. J. Nesbitt. 2006. Review of the Late Triassic dinosaur record from Petrified Forest National Park, Arizona. Museum of Northern Arizona Bulletin 62:160-161.

Sedimentology of Dinosauriform-bearing Triassic Rocks in Poland

I have been away dealing with a personal issue (the sudden passing of my mother), but am taking a much needed break to check messages and such and found this paper in my mailbox (thanks Randy)so I decided to do some therapeutic blogging. It is from last year but provides an important sedimentological and depositional system study of Triassic rocks in Poland which contain the Krasiejow fauna, which is famous for the presence of the dinosauriform Silesaurus opolensis. One of the key aspects of this paper is the support of a Norian age for this fauna rather than a Carnian age as previously supposed. This study removes another "Carnian" record from the record of dinosauriforms demonstrating a global survival of non-dinosaurian dinosauriforms into the Norian (at least in Laurasia)where they were contemporaneous with dinosaurs. This paper also provides evidence that the Krasiejow environment was subtropical and wet, but punctuated with dry periods, thus the climate was monsoonal as has been proposed for the Chinle Formation.

Note: Someone e-mailed me asking about the recent Chinese Jurassic theropod paper. I accidentally deleted your message so please resend.

Gruszka, B. and T. Zielinski. 2008. Evidence for a very low-energy fluvial system: a case study from the dinosaur-bearing Upper Triassic rocks of Southern Poland. Geological Quarterly 52:239-252.

The Upper Triassic succession in S Poland in which dinosaur bones have been found consists predominantly of siltstones and claystones. Three units are distinguished. The lowermost and the upper most units reflect an alluvial environment, whereas the middle one represents lacustrine facies. The lower alluvial unit is interpreted as a record of ephemeral, sinuous, suspended-load channels with rapid vertical accretion. Channel barforms are lacking. The environment is interpreted as a low-energy anastomosing fluvial system. The clayey middle unit is interpreted as having formed in a wide long-lived lake. The top of the lacustrine deposits shows signs of vertisol-type pedogenesis, most probably under subtropical conditions, with seasonally-in duced wet and dry intervals. The upper unit reflects a low-energy meandering river system. Silty point bars were abundant and the channels migrated freely. The energy level of this fluvial system was slightly higher than that of the earlier one, which is interpreted as an effect of base-level lowering in combination with an increasingly humid climate. The almost exclusively silty/clayey alluvial deposits represent an exceptionally rare facies. The drainage basin must have been an extremely flat lowland. The presence of vertebrate bones within the anastomosing and meandering river deposits indicates that low-energy alluvial plains were apparently favourable habitats for both reptiles and amphibians during the Late Triassic: under the subtropical, seasonally dry conditions, the animals must have preferred moist low areas, i.e. the flood basins and abandoned channels on the flat valley floors.

Chinle Silesaurid and the Importance of Field Notes

Adam Yates nailed it. The specimen is the proximal end of the right femur of a silesaurid (PEFO 34347) from the Upper Triassic Blue Mesa Member (Chinle Formation) of Arizona. This specimen is significant because it represents the only known unambiguous silesaurid element from the lower portion of the Chinle Formation and from Arizona. It demonstrates that silesaurids were a portion of the fauna at Petrified Forest National Park.

This specimen was discussed in more detail by Parker et al. (2006) and Nesbitt et al. (2007). It is identical to the proximal ends of the femora of Silesaurus opolensis (Carnian of Poland) and Eucoelophysis baldwini (Norian of New Mexico) and thus cannot be assigned to a specific genus although the age (Norian) and stratigraphic position (Chinle Formation) of the specimen would suggests that it is probably could represent Eucoelophysis rather than Silesaurus. However, until more material is found this cannot be considered. A key characteristic of the proximal end of the femur in Silesaurus, Eucoelophysis, and PEFO 34347 is that the element is triangular in proximal view and has a mediolaterally trending sulcus. Whereas this sulcus is present in other taxa, most notably the pseudosuchian Shuvosaurus, the femur of silesaurids differs in having a subrectangular femoral head in lateral view with a slightly offset head as in dinosaurs.

Now for the promised “interesting” (and frustrating) story regarding this specimen. This specimen was collected sometime in the late 1990s by an unknown individual who was part of a larger research project. It went unrecognized and was included in a large amount of material deemed unworthy of study and potentially to be disposed of. When going through this material to see if anything was salvageable I came across this specimen. Unfortunately, the exact spot where the specimen was collected was not recorded. There are no known field notes for the project and field tags contain minimal information, in this case just a vague geographical reference. This reference is enough to pinpoint the specimen to a small geographical area and limited stratigraphic level; however, it will be nearly impossible to find the rest of the specimen if it exists (and the break is clean, suggesting that more of the specimen was preserved and awaits discovery).

I cannot emphasize enough (and I stress this to, and require it from, all of my employees and interns) the importance of collecting and recording accurate field data on every specimen collected whether the specimen seems important or not. I don’t know how I would function without my past field notes when it comes to identifying and interpreting specific specimens. I have found when I get lax (because of time, weather, arrogance, etc..) I usually end of regretting not having a key piece of information regarding a specimen at some point. At the absolute minimum for EVERY specimen collected there should be GPS coordinates, a photograph of the site showing the surrounding landscape (most important), and a brief description of the sediments and stratigraphic position. Taphonomic notes are also extremely important and often forgotten. Bottom line, you cannot collect too much information. Anyone who has worked with older specimens when simply a stratigraphic unit and state were deemed sufficient information will understand. PEFO 34347 currently represents to earliest known silesaurid from North America, yet its provenance cannot be precisely determined and clarification regarding this specimen depends exclusively on luck. Was more of the specimen preserved? Is it still present and will we be able to stumble across it? I certainly hope so.

REFERENCES

Nesbitt, S.J., Irmis, R.B., and W.G. Parker. 2007. A critical re-evaluation of the Late Triassic dinosaur taxa of North America. Journal of Systematic Palaeontology 5:209-243.

Parker, W.G., Irmis, R.B., and S.J. Nesbitt. 2006. Review of the Late Triassic dinosaur record from Petrified Forest National Park, Arizona. Museum of Northern Arizona Bulletin 62:160-161.