High Latitute Triassic Flora from Antarctica

Escapa, I. H., Taylor, E. L., Cúneo, R., Bomfleur, B., Bergene, J., Serbet, R., and  T. N. Taylor. 2011. Triassic floras of Antarctica: plant diversity and distribution in high paleolatitude communities. Palaios 26:522-544.

Abstract - Continental Triassic sequences in Antarctica are among the most continuous and best represented in Gondwana. Triassic fossil plants have been collected sporadically from Antarctica since the beginning of the twentieth century, but our knowledge of the vegetation during this time has dramatically increased during the last three decades. Here we review the fossil record of Triassic plants as representatives of natural groups from sites along the Transantarctic Mountains, using the fossils as evidence for successive vegetational changes through the Triassic, taking into account that these plant communities were living under particular high-latitude (70° or higher) paleoclimatological conditions, including a polar light regime. Even though our knowledge of the Triassic floras of Antarctica is still incomplete, this survey shows that these floras were remarkably diverse. Lycopsids, equisetaleans, ferns, seed ferns, ginkgoaleans, and conifers were major components of the landscape in Antarctica during this time. The diversity of gymnosperms is exceptional, with almost every major clade of seed plants present, despite the high paleolatitude; however, each clade is often represented by only one or a few genera. The occurrence of permineralized peat, along with compression-impression floras, has increased our knowledge of the morphology, reproductive biology, and evolution of many of the plants in these floras. In general, floral changes in Antarctica during the Triassic can be recognized elsewhere in Gondwana, especially in South America, although a strict correlation based on macrofossils is still not possible. Thus, this contribution represents the first attempt to bring together information on Triassic floras from continental Antarctica (excluding the Antarctic Peninsula) within a biostratigraphic framework and thereby to compare these floras with those from lower latitudes.

Why We Don't Find Cynodonts in the Chinle Formation

Whiteside, J. H., Grogan, D. S., Olsen, P. E., and D. V. Kent. 2011. Climatically driven biogeographic provinces of Late Triassic tropical Pangea. PNAS Published online before print May 13, 2011,
doi: 10.1073/pnas.1102473108  [supplemental info] 

Abstract - Although continents were coalesced into the single landmass Pangea, Late Triassic terrestrial tetrapod assemblages are surprisingly provincial. In eastern North America, we show that assemblages dominated by traversodont cynodonts are restricted to a humid 6° equatorial swath that persisted for over 20 million years characterized by “semiprecessional” (approximately 10,000-y) climatic fluctuations reflected in stable carbon isotopes and sedimentary facies in lacustrine strata. More arid regions from 5–20°N preserve procolophonid-dominated faunal assemblages associated with a much stronger expression of approximately 20,000-y climatic cycles. In the absence of geographic barriers, we hypothesize that these variations in the climatic expression of astronomical forcing produced latitudinal climatic zones that sorted terrestrial vertebrate taxa, perhaps by excretory physiology, into distinct biogeographic provinces tracking latitude, not geographic position, as the proto-North American plate translated northward. Although the early Mesozoic is usually assumed to be characterized by globally distributed land animal communities due to of a lack of geographic barriers, strong provinciality was actually the norm, and nearly global communities were present only after times of massive ecological disruptions.

Stratigraphic Correlations of Gondwanan Basins in India and the "Laurasian" Fauna of Northern Gondwana in the Late Triassic

Here is a fairly obscure, but interesting new paper providing updated correlations of strata (including that of Triassic age) in the Gondwanan Basins of India. The final paper could have benefited from some more editing, but overall it provides a decent synthesis and outlines some of the problems of correlations in this are of Gondwana. Most significant is that there are no absolute time constraints available for these strata because of the lack of datable volcanic materials. The overall climate for the entire Triassic for this area is given as arid with a key sea level regression in the Norian. Note, however, that the authors explicitly use the 2004 ISC timescale, and that under currently proposed revisions to this timescale the Carnian is more restricted.

The Triassic vertebrate fauna of India has always struck me as bizarre, mainly because the Early Triassic fauna is extremely similar to that of the Lystrosaurus assemblage zone of South Africa, and does contain several species of Lystrosaurus. Yet the Middle and Late Triassic fauna has much more of a Laurasian feel to it, especially the Late Triassic fauna, which includes basal phytosaurs such as Parasuchus, and more advanced phytosaurs such as a Leptosuchus-like form. Aetosaurs are represented by a paratypothoracisine form, which are only known from North America and Europe. Finally, metoposaurid temnospondyls are also common as in North America and Europe.

This seems very odd to me given that in paleogeographic reconstructions throughout the Triassic, India is firmly nestled on the western edge of modern day Africa, contacting Australia, and Antarctica, and separated from North America by all of northern Africa and from Europe by the Tethys. Unfortunately the Triassic fauna of North Africa is extremely sparse, although the Moroccan fauna is very similar to that of India and Laurasia. Perhaps Northern Africa served as a corridor for the Laurasian-type fauna, whereas there was some restriction from South America and South Africa in the Latest Triassic.  Thus it seems that at this time there are not distinct Laurasian and Gondwanan distributions of tetrapods, but rather separate southern Gondwana and northern Gondwana/Laurasia distributions.

Mukhopadhyay, G., Mukhopadhyay, S. K., Roychowdhury, M., and P. K. Parui. 2010. Stratigraphic Correlation between Different Gondwana Basins of India.
Journal of the Geological Society of India 76:251-266. DOI: 10.1007/s12594-010-0097-6

Abstract - Gondwana Basins of India occur within the suture zones of Precambrian cratonic blocks of Peninsular India along some linear belts. More than 99% of the total coal resource of the country is present within these basins. The basins are demarcated by boundary faults having graben or half-graben geometry. These basins preserve a thick sedimentary pile deposited over nearly 200 million years from latest Carboniferous to Lower Cretaceous. However, due to lack of well-constrained data, age of most of the formations is assigned tentatively. This has resulted in diversified views on both intra- and inter-basinal stratigraphic correlation particularly in case of Upper Gondwana formations. It is well recognised that there are distinct spatial and temporal similarities in lithological, faunal and floral distribution in different Gondwana Basins of southern continents, including India, that were once part of supercontinent Gondwanaland. To address the problems of Indian Gondwana stratigraphy, during the present study, some unique events, also recognised in other parts of Gondwanaland, like marine flooding surfaces, large scale tectonic events or major change in depositional environment have been used as a tool for temporal correlation within the Gondwana Basins of India. Many of these events have been dated from different basins elsewhere. Considering these major events as time planes the total time span of deposition in Gondwana Basins has been classified into seven time slots. Recognition of these time planes helps in interbasinal correlation of different formations in Indian Gondwana basins and assigning the age, wherever available. This approach also helps in better understanding of basinal history. Unless otherwise mentioned, the time scale proposed by International Commission on Stratigraphy (2004) has been followed in this paper.

Biogeography of Triassic tetrapods

Interesting approach and personally timely as I just happen to really be into Triassic biogeography at the moment; however, I think this study suffers somewhat from the usual culprits, lack of taxon sampling and unresolved taxonomic issues (weak primary data).  For example, basal phytosaurs are completely left out which have an early and widespread appearance in the Late Triassic.  These are extremely important as they are found in Europe, North America, North Africa, and interestingly in India.  Including these would have changed several of the paper's conclusions.  Likewise, the aetosaur Calyptosuchus (Stagonolepis) wellesi is purposefully left out as is Coahomasuchus, which are two important datapoints for aetosaurs in the Ischigualastian of North America.  I wish I had been contacted regarding this study as I would have gladly suggested some less "key" aetosaur taxa to leave out if better resolution was needed for the basal aetosaurines.  It is also not entirely clear if the Ischigulastian and Coloradian are applicable outside of South America.  It appears that they are presented as equivalent to the Otischalkian + Adamanian and Revueltian + Apachean of North America.  This was first put forth by Max Langer in 2005 and has some support, but needs further testing, especially regarding the relationships of all of the Stagonolepis-like aetosaurs, the lack of good phytosaur material in South America, and lack of good (or any) comparable rhynchosaur, cynodont, and sauropodomorph material in North America.

On a side note, it was nice to see Heliocanthus used in there.

Ezcurra, M. D. 2010. Biogeography of Triassic tetrapods: evidence for provincialism and driven sympatric cladogenesis in the early evolution of modern tetrapod lineages. Proceedings of the Royal Society B, doi:10.1098/rspb.2010.0508 Published online.

Abstract - Triassic tetrapods are of key importance in understanding their evolutionary history, because several tetrapod clades, including most of their modern lineages, first appeared or experienced their initial evolutionary radiation during this Period. In order to test previous palaeobiogeographical hypotheses of Triassic tetrapod faunas, tree reconciliation analyses (TRA) were performed with the aim of recovering biogeographical patterns based on phylogenetic signals provided by a composite tree of Middle and Late Triassic tetrapods. The TRA found significant evidence for the presence of different palaeobiogeographical patterns during the analysed time spans. First, a Pangaean distribution is observed during the Middle Triassic, in which several cosmopolitan tetrapod groups are found. During the early Late Triassic a strongly palaeolatitudinally influenced pattern is recovered, with some tetrapod lineages restricted to palaeolatitudinal belts. During the latest Triassic, Gondwanan territories were more closely related to each other than to Laurasian ones, with a distinct tetrapod fauna at low palaeolatitudes. Finally, more than 75 per cent of the cladogenetic events recorded in the tetrapod phylogeny occurred as sympatric splits or within-area vicariance, indicating that evolutionary processes at the regional level were the main drivers in the radiation of Middle and Late Triassic tetrapods and the early evolution of several modern tetrapod lineages.