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Galton, P. M., and D. Kermack. 2010. The anatomy of Pantydraco caducus, a very basal sauropodomorph dinosaur from the Rhaetian (Upper Triassic) of South Wales, UK. Revue de Paléobiologie, Genève 29 (2) : 341-404.
Abstract - Apart from characters of the dentary (proportional shortness, also Saturnalia tupiniquim) and vertebrae of proximal third of tail (posterior position of antero-posteriorly short neural spine on arch, absence of ventral furrowing on centra ; both also Efraasia minor), the bones of very basal sauropodomorphs from the Rhaetian (Upper Triassic) fissure fill in Clifton near Bristol (including holotypes of Thecodontosaurus antiquus, Asylosaurus yalensis ; taxon with robust humerus) share no diagnostic characters with those of Pantydraco caducus from the Rhaetian of South Wales. This taxon is diagnosed by four autapomorphies : median fusion of the nasals, prominent medial tubercle from posterior part of surangular, epipophyses of cervical vertebrae 3-5 form flat plates that overhang the posterior margins of the postzygapophyseal facets but do not form raised ridges on the dorsal surface of the postzygapophysis, and the fossae, possibly pneumatic, prominent on the neurocentral suture of cervical vertebrae 6-8 and shallow on the lateral part of centrum 9. The dentary is deep and short, with maximum height of more than 20 % of length (also Saturnalia tupiniquim, Thecodontosaurus antiquus), occupying less than 40 % of total mandibular length (also Saturnalia tupiniquim). The ilium is tall with a long anterior process (also Anchisaurus polyzelus) and a short postacetabular process with the plesiomorphic absence of a brevis shelf. Other plesiomorphic characters include the lack of a buccal emargination on the dentary (also Saturnalia tupiniquim); teeth all recurved in lateral view; neck short with mid-cervical centra less than three times as long as wide (also Riojasaurus incertus, “Gyposaurus” sinensis) ; and humerus with a uniquely small tubercle medial to head and an antero-posteriorly low (also Asylosaurus yalensis) asymmetrical deltopectoral crest with the apex at 40 % of humeral length. At least three different sized individuals are preserved and the new skeletal reconstruction, with the posterior region scaled up to match the larger holotype, indicates a more bipedal animal than the more quadrupedal pose based on the incorrect premise that the two main blocks represent one individual. The specimens represent juveniles as indicated by the proportionally large skull with large orbits and short, high snout, slender postorbital bone, separation of most of bones of braincase, large metotic fissure and fenestra ovalis, low maxillary and dentary tooth counts, open neurocentral sutures of cervical vertebrae, and incomplete ossification of distal ends of the femur and metatarsals. The diversity of basal sauropodomorphs from the Rhaetian of Wales and England adjacent to the Severn Estuary is high with five taxa (Pantydraco caducus, three from Clifton, large basal sauropod Camelotia borealis from Somerset).
Showing posts with label Rhaetian. Show all posts
Showing posts with label Rhaetian. Show all posts
More Evidence for Long Norian and Rhaetian Stages
Carnian-Norian boundary: ~226.6 Ma;
Lacian-Alaunian boundary: ~216.4 Ma (fairly equivalent to the old Carnian/Norian boundary);
Alaunian-Sevatian 1 boundary: 211.8 Ma;
Norian Rhaetian boundary: ~209.8 Ma.
Thus according to these data the early Norian is approximately 10 million years long, the middle Norian is 4.5 million years old, and the late Norian is 2 million years old for an approximately 17 million year long Norian stage. The Rhaetian is approximately 9 million years long.
Proposed biostratigraphic data from palynomorphs and conchostracans suggesting a shorter Norian are refuted.
Hüsing, S. K.,
Abstract - The Global Stratotype Section and Point (GSSP) for the Rhaetian Stage has recently been proposed at Steinbergkogel in Austria. We re-sampled the Steinbergkogel sections (STK-A and STK-B + C) in high-resolution to establish a robust magnetostratigraphy that allows global correlation. The palaeomagnetic signal at Steinbergkogel is composed of three components, which can be separated by thermal demagnetization. The highest temperature component, revealed between 280/300 and maximum 600 °C is of dual polarity and is interpreted as primary. Rock magnetic experiments showed that the signal is carried by magnetite. Our results allow correlation between the two individual Steinbergkogel outcrops. Subsequently, we correlate the two key biostratigraphic horizons for the base of the Rhaetian, the FO of M. hernsteini and the FAD of M. posthernsteini to other sections of the Tethys domain. The correlation to the astronomically dated continental successions of the Newark basin indicates that these positions for the base of the Rhaetian are most likely determined in chrons E16n and E16r, respectively. This correlation is confirmed by cyclostratigraphic control on the marine Pizzo Mondello (Italy) section, where a combination of long period Milankovitch cycles (~ 175-Myr) and short-eccentricity cycles (~ 100-kyr) provide additional correlation constraints, respectively supporting a long duration of the Rhaetian. Our study implies that the Norian and Rhaetian Stages have durations of ~ 17 and ~ 9 Myr.
REFERENCES
Muttoni, G., Kent, D.V., Jadoul, F., Olsen, P.E., Rigo, M., Galli, M.T., and A. Nicora. 2010. Rhaetian magneto-biostratigraphy from the Southern Alps (Italy): constraints on Triassic chronology. Palaeogeography, Palaeoclimatology, Palaeoecology 285: 1–16.
Muttoni, G., Kent, D.V., Olsen, P.E., Di Stefano, P., Lowrie, W., Bernasconi, S.M., and F. M. Hernandez. 2004. Tethyan magnetostratigraphy from Pizzo Mondello (Sicily)and correlation to the Late Triassic Newark astrochronological polarity time scale. Geological Society of America Bulletin 116: 1043–1058.
Rhaetian Ichnology and Geochemistry of the Westbury Formation, England
Allington-Jones, L., Braddy, S. J., and Trueman, C. N. 2010. Palaeoenvironmental implications of the ichnology and geochemistry of the Westbury Formation (Rhaetian), Westburyon- Severn, South-West England. Palaeontology 53:491-506. doi: 10.1111/j.1475-4983.2010.00947.x
Abstract - The Westbury Formation (Rhaetian) beds of Westbury Garden Cliff, Westbury-on-Severn, west of Gloucester, Britain, show an unusual combination of features. Both deep water and emergent characteristics are present within the sediments and the trace fossils. The ichnoassemblage consists of abundant Selenichnites, Planolites beverlyensis and Lockeia with rarer Oniscoidichnus, Chondrites, Rhizocorallium irregulare, Taenidium serpentium, an unusual form of Walcottia and Merostomichnites-like traces. These trace fossils display an interesting relationship with the sediments: low-energy Cruziana ichnofacies is found within high-energy sandstones. The sandstones are interbedded with laminated mudstones, apparently deposited in deep water, but some aspects of the ichnoassemblage, preservation and sedimentation indicate shallower water. One new trace fossil, Radichnus allingtona igen. et isp. nov., closely resembles the traces of modern fiddler crabs and imply emergence, by analogy. This ichnofauna is similar to early stage disaster colonisation in recent experiments in Long Island Sound (south of Connecticut, USA) and with storm influenced deposits within the Cardium Formation (Seebe, Alberta, Canada). This indicates a lagoonal environment with influxes of sand and oxygen. Total organic carbon levels were found to fluctuate greatly between stratigraphic layers but remained relatively high. This implies low oxygen conditions. The abundance of sulphur (in pyrite) also supports an interpretation of anoxic conditions, and low sedimentation rates within the shale layers. A restricted shallow basin or lagoonal environment is proposed for the palaeoenvironment, with fluctuating oxygen influencing diversity.
Abstract - The Westbury Formation (Rhaetian) beds of Westbury Garden Cliff, Westbury-on-Severn, west of Gloucester, Britain, show an unusual combination of features. Both deep water and emergent characteristics are present within the sediments and the trace fossils. The ichnoassemblage consists of abundant Selenichnites, Planolites beverlyensis and Lockeia with rarer Oniscoidichnus, Chondrites, Rhizocorallium irregulare, Taenidium serpentium, an unusual form of Walcottia and Merostomichnites-like traces. These trace fossils display an interesting relationship with the sediments: low-energy Cruziana ichnofacies is found within high-energy sandstones. The sandstones are interbedded with laminated mudstones, apparently deposited in deep water, but some aspects of the ichnoassemblage, preservation and sedimentation indicate shallower water. One new trace fossil, Radichnus allingtona igen. et isp. nov., closely resembles the traces of modern fiddler crabs and imply emergence, by analogy. This ichnofauna is similar to early stage disaster colonisation in recent experiments in Long Island Sound (south of Connecticut, USA) and with storm influenced deposits within the Cardium Formation (Seebe, Alberta, Canada). This indicates a lagoonal environment with influxes of sand and oxygen. Total organic carbon levels were found to fluctuate greatly between stratigraphic layers but remained relatively high. This implies low oxygen conditions. The abundance of sulphur (in pyrite) also supports an interpretation of anoxic conditions, and low sedimentation rates within the shale layers. A restricted shallow basin or lagoonal environment is proposed for the palaeoenvironment, with fluctuating oxygen influencing diversity.
New Biostratigraphical Constraints for the Norian⁄Rhaetian Boundary
Once even questioned as a valid stage the Rhaetian has made quite a comeback research-wise, especially since the GSSP for the Norian/Rhaetian boundary has yet to be determined.
Giordano, N., Rigo, M., Ciarapica G. & Bertinelli A. 2010. New biostratigraphical constraints for the Norian ⁄Rhaetian boundary: data from Lagonegro Basin, Southern Apennines, Italy. Lethaia, 10.1111/j.1502-3931.2010.00219.x.
Abstract - Four stratigraphic sections belonging to Lagonegro succession (Southern Apennines) at Mt S. Enoc, Pignola-Abriola, Sasso di Castalda and Mt Volturino have been studied in detail under to provide a new micro-palaeontological data set based on conodonts and radiolarians for the characterization of the Norian ⁄Rhaetian interval. The studied sections represent the different settings of the Lagonegro Basin (from proximal to distal facies) and permit a detailed, integrated, biostratigraphy of the Calcari con Selce (cherty limestones) and Scisti Silicei formations (bedded cherts with radiolarians) to be drawn up. The upper portion of the Calcari con Selce Formation, exhibits intermediate characteristics between the Calcari con Selce and Scisti Silicei Formation, in particular the progressive decrease in carbonate content against an increase in shales and cherts. Within the four sections studied, the Norian ⁄Rhaetian interval has been documented both with conodonts and radiolarians. Because of the continuity and the absence of condensed facies, it has been possible to recognize the morphocline between species Misikella hernsteini and Misikella posthernsteini, here represented by all the transitional forms characterized by common features between the two species, gathered in three evolutionary steps. Moreover, the morphocline between M. hernsteini and M. posthernsteini has been involved in the definition of the Norian ⁄ Rhaetian Boundary, recognizing thus the FAD of M. posthernsteini, one of the possible biomarkers proposed for the boundary. The rich, well-preserved, radiolarian associations of Pignola-Abriola, Sasso di Castalda and Mt Volturino permit the correlation of Tethyan and American conodont successions, highlighting the importance of the mostly coincident occurrences of M. posthernsteini and Epigondolella mosheri morphotype A, which correspond to the base of Proparvicingula moniliformis A. Z. and the disappearance of bivalve Monotis. These coincident bioevents are used here to define the base of the Rhaetian stage.
Giordano, N., Rigo, M., Ciarapica G. & Bertinelli A. 2010. New biostratigraphical constraints for the Norian ⁄Rhaetian boundary: data from Lagonegro Basin, Southern Apennines, Italy. Lethaia, 10.1111/j.1502-3931.2010.00219.x.
Abstract - Four stratigraphic sections belonging to Lagonegro succession (Southern Apennines) at Mt S. Enoc, Pignola-Abriola, Sasso di Castalda and Mt Volturino have been studied in detail under to provide a new micro-palaeontological data set based on conodonts and radiolarians for the characterization of the Norian ⁄Rhaetian interval. The studied sections represent the different settings of the Lagonegro Basin (from proximal to distal facies) and permit a detailed, integrated, biostratigraphy of the Calcari con Selce (cherty limestones) and Scisti Silicei formations (bedded cherts with radiolarians) to be drawn up. The upper portion of the Calcari con Selce Formation, exhibits intermediate characteristics between the Calcari con Selce and Scisti Silicei Formation, in particular the progressive decrease in carbonate content against an increase in shales and cherts. Within the four sections studied, the Norian ⁄Rhaetian interval has been documented both with conodonts and radiolarians. Because of the continuity and the absence of condensed facies, it has been possible to recognize the morphocline between species Misikella hernsteini and Misikella posthernsteini, here represented by all the transitional forms characterized by common features between the two species, gathered in three evolutionary steps. Moreover, the morphocline between M. hernsteini and M. posthernsteini has been involved in the definition of the Norian ⁄ Rhaetian Boundary, recognizing thus the FAD of M. posthernsteini, one of the possible biomarkers proposed for the boundary. The rich, well-preserved, radiolarian associations of Pignola-Abriola, Sasso di Castalda and Mt Volturino permit the correlation of Tethyan and American conodont successions, highlighting the importance of the mostly coincident occurrences of M. posthernsteini and Epigondolella mosheri morphotype A, which correspond to the base of Proparvicingula moniliformis A. Z. and the disappearance of bivalve Monotis. These coincident bioevents are used here to define the base of the Rhaetian stage.
Rhaetian Magneto-biostratigraphy from the Southern Alps (Italy): Constraints on Triassic Chronology
This is fairly significant as it argues for quite a long Rhaetian Stage of the latest Triassic. If this is correct it means that a sizable portion of the upper part of the Chinle Formation could actually be Rhaetian in age. Not bad considering not too long ago workers were arguing if the Rhaetian was really even applicable outside of the marine realm.
Muttoni, G., Kent, D.V., Jadoul, F., Olsen, P.E., Rigo, M., Galli, M.T., and A. Nicora. 2009. Rhaetian magneto-biostratigraphy from the Southern Alps (Italy): Constraints on Triassic chronology. Palaeogeography, Palaeoclimatology, Palaeoecology. doi: 10.1016/j.palaeo.2009.10.014.
ABSTRACT: New Late Triassic-earliest Jurassic magneto-biostratigraphic data have been obtained from three overlapping sections in the Southern Alps, Italy (Costa Imagna, Brumano, Italcementi Quarry), comprised of ~ 520 m of shallow marine carbonates outcropping in stratigraphic continuity. Characteristic magnetic components of presumed depositional age record a sequence of 9 normal and reverse polarity magnetozones (as defined by at least three stratigraphically superposed samples) linked by conodont and palynofloral evidence from this study and the literature to Rhaetian to Triassic–Jurassic boundary age. This represents a significantly larger number of polarity zones than previously recognized in more condensed
Rhaetian sections from the literature, and by inference represents more time. These data are placed in a broader Late Triassic temporal framework by means of correlations to published magneto-biostratigraphic data from the Tethyan marine Pizzo Mondello section and the Newark astronomical polarity time scale (APTS). This framework is consistent with a position of the
Norian–Rhaetian boundary (as defined at Brumano and Pizzo Mondello by the first appearance of Misikella posthernsteini) within Newark magnetozones E17r–E19r in the ~ 207–210 Ma time interval, in basic agreement with the position originally estimated in the Newark using pollen biostratigraphy (E18 at 208–209 Ma). This framework is also consistent with the position of
the Triassic–Jurassic boundary interval (placed at Italcementi Quarry at the acme of Kraeuselisporites reissingeri coincident with a negative carbon isotope excursion) correlative to just above Newark magnetozone E23r and just below the oldest CAMP lavas dated at ~ 202 Ma. Hence, we estimate the duration of the Rhaetian to be ~ 5.5–8.5 Myr (or even longer if the
Triassic–Jurassic boundary is instead placed above the negative carbon isotope excursion as at Kuhjoch, which is the designated GSSP for the base of the Hettangian), and encompassing 9 magnetozones. This duration contrasts with a duration of ~ 2 Myr and only ~ 4 magnetozones in several alternative published magneto-biostratigraphic schemes.
Muttoni, G., Kent, D.V., Jadoul, F., Olsen, P.E., Rigo, M., Galli, M.T., and A. Nicora. 2009. Rhaetian magneto-biostratigraphy from the Southern Alps (Italy): Constraints on Triassic chronology. Palaeogeography, Palaeoclimatology, Palaeoecology. doi: 10.1016/j.palaeo.2009.10.014.
ABSTRACT: New Late Triassic-earliest Jurassic magneto-biostratigraphic data have been obtained from three overlapping sections in the Southern Alps, Italy (Costa Imagna, Brumano, Italcementi Quarry), comprised of ~ 520 m of shallow marine carbonates outcropping in stratigraphic continuity. Characteristic magnetic components of presumed depositional age record a sequence of 9 normal and reverse polarity magnetozones (as defined by at least three stratigraphically superposed samples) linked by conodont and palynofloral evidence from this study and the literature to Rhaetian to Triassic–Jurassic boundary age. This represents a significantly larger number of polarity zones than previously recognized in more condensed
Rhaetian sections from the literature, and by inference represents more time. These data are placed in a broader Late Triassic temporal framework by means of correlations to published magneto-biostratigraphic data from the Tethyan marine Pizzo Mondello section and the Newark astronomical polarity time scale (APTS). This framework is consistent with a position of the
Norian–Rhaetian boundary (as defined at Brumano and Pizzo Mondello by the first appearance of Misikella posthernsteini) within Newark magnetozones E17r–E19r in the ~ 207–210 Ma time interval, in basic agreement with the position originally estimated in the Newark using pollen biostratigraphy (E18 at 208–209 Ma). This framework is also consistent with the position of
the Triassic–Jurassic boundary interval (placed at Italcementi Quarry at the acme of Kraeuselisporites reissingeri coincident with a negative carbon isotope excursion) correlative to just above Newark magnetozone E23r and just below the oldest CAMP lavas dated at ~ 202 Ma. Hence, we estimate the duration of the Rhaetian to be ~ 5.5–8.5 Myr (or even longer if the
Triassic–Jurassic boundary is instead placed above the negative carbon isotope excursion as at Kuhjoch, which is the designated GSSP for the base of the Hettangian), and encompassing 9 magnetozones. This duration contrasts with a duration of ~ 2 Myr and only ~ 4 magnetozones in several alternative published magneto-biostratigraphic schemes.
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